Umbrous is a a dominant allele which causes a dark spine. You usually see it on sable. It also shows up on agouti-based varieties (agouti, argente, cinnamon) where it is a fault. It doesn't show up on most self varieties, and even on agouti, argente, and cinnamon it's dominance is modified by homozygosity of the A-locus (which is to say that a mouse who is A/A U/* or a/a U/* won't have a stripe or won't have as dark a stripe as one who is A/a U/*), so you don't always see it. Mice in the same litter can be umbrous and only those who are heterozygous on the A locus (A/a U/*) will show umbrous. Once it's present it's very hard to get rid of!